T4SS ID	16
Strain	Agrobacterium rhizogenes plasmid pRi2659 (NC_010841)
Replicon	plasmid pRi2659 [Browse all T4SS(s) in this replicon]
Accession	NC_010841
Location	157955..177542
Name	VirB
Function	effector translocation and conjugation
Classification	Type IVA; Type P
This T4SS has been mentioned in a genome sequencing report.

T4SS components

Component	VirB1	VirB2	VirB3	VirB4	VirB5	VirB6	VirB7	VirB8	VirB9
Number	1	1	1	1	1	1	1	1	1
Component	VirB10	VirB11	VirD4
Number	1	1	1

The information of T4SS components from NC_010841
#	Locus tag (Gene)	Coordinates [+/-], size (bp)	Protein GI	Product	Component
1	pRi2659_p120	153072..154424 [+], 1353	190404463	rcorf119
2	pRi2659_p121	154628..157183 [+], 2556	190404464	two-component VirA-like sensor kinase
3	pRi2659_p122	157955..158677 [+], 723	190404465	type IV secretion system lytic transglycosylase VirB1	VirB1
4	pRi2659_p123	158677..159042 [+], 366	190404466	type IV secretion system pilin subunit VirB2	VirB2
5	pRi2659_p124	159039..159365 [+], 327	190404467	type IV secretion system protein VirB3	VirB3
6	pRi2659_p125	159365..161734 [+], 2370	190404468	type IV secretion system protein VirB4	VirB4
7	pRi2659_p126	161751..162419 [+], 669	190404469	type IV secretion system protein VirB5	VirB5
8	pRi2659_p127	162486..163373 [+], 888	190404470	type IV secretion system protein VirB6	VirB6
9	pRi2659_p128	163401..163568 [+], 168	190404471	type IV secretion system lipoprotein VirB7	VirB7
10	pRi2659_p129	163555..164268 [+], 714	190404472	type IV secretion system protein VirB8	VirB8
11	pRi2659_p130	164265..165143 [+], 879	190404473	type IV secretion system protein VirB9	VirB9
12	pRi2659_p131	165143..166273 [+], 1131	190404474	type IV secretion system protein VirB10	VirB10
13	pRi2659_p132	166307..167341 [+], 1035	190404475	type IV secretion system protein VirB11	VirB11
14	pRi2659_p133	167577..168302 [+], 726	228861208	two-component response regulator VirG
15	pRi2659_p134	168409..168765 [-], 357	190404477	rcorf133
16	pRi2659_p135	168741..169106 [-], 366	190404478	rcorf134
17	pRi2659_p136	169270..169875 [-], 606	190404479	putative crown gall tumor protein VirC2
18	pRi2659_p137	169878..170573 [-], 696	190404480	putative crown gall tumor protein VirC1
19	pRi2659_p138	170828..171274 [+], 447	190404481	type IV secretion system T-DNA border endonuclease VirD1
20	pRi2659_p139	171300..172646 [+], 1347	190404482	type IV secretion system T-DNA border endonuclease VirD2
21	pRi2659_p140	172477..175500 [+], 3024	190404483	rcorf139
22	pRi2659_p141	175560..177542 [+], 1983	190404484	type IV secretion system protein VirD4	VirD4
23	pRi2659_p142	177650..180157 [+], 2508	190404485	rcorf141
24	pRi2659_p143	180410..181339 [+], 930	190404486	rcorf142

Genes in the 5-Kb flanking regions if available, or non-essential genes in the T4SS gene cluster if any.

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Proteins        Genes

This system delivers both DNA and proteins into plant cell. The transferred DNA (T-DNA) region: NC_010841: 1..4247)

Effectors

GALLS, VirF, VirD2, VirD5, VirE3

The information of protein effectors
#	Locus tag (Gene)	Coordinates [+/-], size (bp)	Protein GI	Product	*
1	pRi2659_p029	34384..39693 [-], 5310	190404372	rcorf28	GALLS
2	pRi2659_p115	146513..147466 [+], 954	190404458	rcorf114	VirF
3	pRi2659_p139	171300..172646 [+], 1347	190404482	type IV secretion system T-DNA border endonuclease VirD2	VirD2
4	pRi2659_p142	177650..180157 [+], 2508	190404485	rcorf141	VirD5
5	pRi2659_p144	181443..182798 [+], 1356	190404487	rcorf143	VirE3

 T4SE derived from experimental literature.

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Proteins        Genes

(1) Hodges LD; Lee LY; McNett H; Gelvin SB; Ream W (2009). The Agrobacterium rhizogenes GALLS gene encodes two secreted proteins required for genetic transformation of plants. J Bacteriol. 191(1):355-64. [PudMed:18952790]

This literature contains experimental investigation