NZ_CP0514311..217533

MGE detailed information

MGE type: Non-Mobilizable plasmid        MGE length: 217533 bp
Accession: NZ_CP051431        Location: 1..217533         Organism: Escherichia sp. SCLE84       Replicon: plasmid pSCLE1


Gene structure


The 5 kb flanking regions of this MGE are displayed.

Locus tag Coordinates Strand Size (bp) Protein ID Product Description
HGD79_RS24330 () 1..876 + 876 WP_000594612.1 RepB family plasmid replication initiator protein -
HGD79_RS24335 () 2025..2378 + 354 WP_000423602.1 hypothetical protein -
HGD79_RS24340 () 2396..2746 + 351 WP_024152015.1 plasmid transfer protein -
HGD79_RS24345 () 2758..3546 + 789 WP_000783153.1 pilus assembly protein -
HGD79_RS24350 () 3546..4817 + 1272 WP_000592090.1 type-F conjugative transfer system secretin TraK -
HGD79_RS24355 () 4819..5280 + 462 WP_000521240.1 hypothetical protein -
HGD79_RS24360 () 5270..6625 + 1356 WP_000351841.1 IncHI-type conjugal transfer protein TrhB -
HGD79_RS24365 () 6693..7115 + 423 WP_000251250.1 hypothetical protein -
HGD79_RS24370 () 7234..7986 + 753 WP_183079335.1 protein-disulfide isomerase HtdT -
HGD79_RS24375 () 7996..8946 + 951 WP_001022587.1 IncHI-type conjugal transfer lipoprotein TrhV -
HGD79_RS24380 () 8955..10844 + 1890 Protein_10 TraC family protein -
HGD79_RS24385 () 11015..12214 - 1200 WP_000948429.1 IS91 family transposase ISSbo1 Transposase
HGD79_RS24390 () 12224..12412 - 189 WP_000957857.1 hypothetical protein -
HGD79_RS24395 () 12527..13324 + 798 Protein_13 AAA family ATPase -
HGD79_RS24400 () 17161..18414 + 1254 WP_032084678.1 Plasmid partition protein A VF
HGD79_RS24405 () 18411..19415 + 1005 WP_032192066.1 ParB N-terminal domain-containing protein -
HGD79_RS24410 () 19477..19866 - 390 WP_001572421.1 hypothetical protein -
HGD79_RS24415 () 19863..20384 - 522 WP_168848542.1 hypothetical protein -
HGD79_RS24420 () 20377..21213 - 837 WP_042634275.1 hypothetical protein -
HGD79_RS24425 () 21210..22157 - 948 WP_046788422.1 metallophosphoesterase -
HGD79_RS24430 () 22521..23558 + 1038 WP_000818955.1 hypothetical protein -
HGD79_RS24435 () 23570..24208 + 639 WP_001107537.1 hypothetical protein -
HGD79_RS24440 () 24612..25064 + 453 WP_000952689.1 hypothetical protein -
HGD79_RS24445 () 25054..25479 + 426 WP_000836980.1 hypothetical protein -
HGD79_RS24450 () 25488..26042 + 555 WP_000235777.1 hypothetical protein -
HGD79_RS24455 () 26162..30466 + 4305 WP_001232107.1 IncHI-type conjugal transfer protein RSP -
HGD79_RS24460 () 30744..31256 + 513 WP_001286759.1 IncHI-type conjugal transfer protein TrhF -
HGD79_RS24465 () 31243..32754 + 1512 WP_001256133.1 hypothetical protein -
HGD79_RS24470 () 32899..33954 + 1056 WP_181196458.1 IncHI-type conjugal transfer protein TrhU -
HGD79_RS24475 () 33973..37162 + 3190 Protein_29 IncHI-type conjugal transfer protein TrhN -
HGD79_RS24480 () 37191..38063 - 873 WP_000771832.1 hypothetical protein -
HGD79_RS24485 () 38243..40027 + 1785 WP_042634277.1 IncHI-type conjugal transfer helicase TrhI -
HGD79_RS24490 () 40422..41294 + 873 WP_000041927.1 hypothetical protein -
HGD79_RS24495 () 41352..41729 + 378 WP_000120822.1 hypothetical protein -
HGD79_RS24500 () 41790..42767 + 978 WP_042634278.1 hypothetical protein -
HGD79_RS24505 () 42825..43883 + 1059 WP_000063335.1 AAA family ATPase -
HGD79_RS24510 () 44015..44188 + 174 WP_001176647.1 hypothetical protein -
HGD79_RS24515 () 44252..45379 + 1128 WP_000174395.1 DUF3150 domain-containing protein -
HGD79_RS24520 () 45457..47469 + 2013 WP_046788423.1 VWA domain-containing protein -
HGD79_RS24525 () 47541..47762 + 222 WP_042634307.1 hypothetical protein -
HGD79_RS24530 () 47876..49108 + 1233 WP_000220381.1 toprim domain-containing protein -
HGD79_RS24535 () 49379..49900 + 522 WP_001010933.1 hypothetical protein -
HGD79_RS24540 () 49979..50857 + 879 WP_000762004.1 hypothetical protein -
HGD79_RS24545 () 50928..51863 + 936 WP_000691118.1 hypothetical protein -
HGD79_RS24550 () 51932..52852 + 921 WP_000991699.1 hypothetical protein -
HGD79_RS24555 () 52919..53857 + 939 WP_000472043.1 hypothetical protein -
HGD79_RS24560 () 54029..54985 + 957 WP_046788424.1 hypothetical protein -
HGD79_RS24565 () 55249..55524 + 276 WP_001004201.1 hypothetical protein -
HGD79_RS24570 () 55569..56159 + 591 WP_000578892.1 hypothetical protein -
HGD79_RS24575 () 56167..56424 + 258 WP_001371933.1 hypothetical protein -
HGD79_RS24580 () 56498..57034 + 537 WP_000107168.1 hypothetical protein -
HGD79_RS24585 () 57114..57497 + 384 WP_001424628.1 hypothetical protein -
HGD79_RS24590 () 57588..58550 + 963 WP_000119427.1 DNA replication protein -
HGD79_RS24595 () 58560..59465 + 906 WP_001198594.1 hypothetical protein -
HGD79_RS24600 () 59401..59745 + 345 WP_001805195.1 hypothetical protein -
HGD79_RS24605 () 59767..60942 - 1176 WP_000795947.1 tyrosine-type recombinase/integrase -
HGD79_RS24610 () 61113..61325 + 213 WP_001285422.1 hypothetical protein -
HGD79_RS24615 () 61686..62768 + 1083 WP_001232449.1 hypothetical protein -
HGD79_RS24620 () 62935..64434 - 1500 WP_001284313.1 kinase -
HGD79_RS24625 () 64460..66097 - 1638 WP_000081622.1 protein phosphatase 2C domain-containing protein -
HGD79_RS24630 () 66097..67137 - 1041 WP_001253653.1 VWA domain-containing protein -
HGD79_RS24635 () 67223..67861 - 639 WP_001253658.1 VWA domain-containing protein -
HGD79_RS24640 () 67861..68502 - 642 WP_000116681.1 TerD family protein -
HGD79_RS24645 () 68525..69163 - 639 WP_001388628.1 VWA domain-containing protein -
HGD79_RS24650 () 69626..70093 - 468 WP_001176767.1 tellurium resistance protein TerW -
HGD79_RS24655 () 70111..71319 - 1209 WP_000506887.1 DUF4236 domain-containing protein -
HGD79_RS24660 () 71330..72286 - 957 WP_000797366.1 HpcH/HpaI aldolase/citrate lyase family protein -
HGD79_RS24665 () 72286..73365 - 1080 WP_042634304.1 cysteine protease StiP family protein -
HGD79_RS24670 () 73367..74140 - 774 WP_001040059.1 hypothetical protein -
HGD79_RS24675 () 74133..75275 - 1143 WP_001285478.1 phosphoribosyltransferase domain-containing protein -
HGD79_RS24680 () 75285..76343 - 1059 WP_001035162.1 ATP-grasp domain-containing protein -
HGD79_RS24685 () 76667..77248 + 582 WP_042634303.1 tellurium resistance TerZ family protein -
HGD79_RS24690 () 77248..78405 + 1158 WP_001054787.1 TerD family protein -
HGD79_RS24695 () 78428..78883 + 456 WP_000007448.1 tellurite resistance TerB family protein -
HGD79_RS24700 () 78906..79946 + 1041 WP_000255079.1 tellurium resistance membrane protein TerC -
HGD79_RS24705 () 79995..80573 + 579 WP_000116677.1 tellurium resistance membrane protein TerD -
HGD79_RS24710 () 80641..81216 + 576 WP_000301242.1 TerD family protein -
HGD79_RS24715 () 81645..82886 + 1242 WP_001053338.1 VWA domain-containing protein -
HGD79_RS24720 () 83449..83730 - 282 WP_000077926.1 hypothetical protein -
HGD79_RS24725 () 83780..83971 - 192 WP_000398480.1 hypothetical protein -
HGD79_RS24730 () 84063..84404 - 342 WP_001151575.1 hypothetical protein -
HGD79_RS24735 () 84777..85169 + 393 WP_000341066.1 hypothetical protein -
HGD79_RS24740 () 85773..86066 + 294 WP_024136327.1 helix-turn-helix transcriptional regulator Antitoxin Detail
HGD79_RS24745 () 86071..87396 + 1326 WP_086722013.1 type II toxin-antitoxin system HipA family toxin Toxin Detail
HGD79_RS24750 () 87457..87663 + 207 WP_000134171.1 hypothetical protein -
HGD79_RS24755 () 87765..88175 + 411 WP_000985911.1 hypothetical protein -
HGD79_RS24760 () 88188..89003 + 816 WP_001572439.1 HNH endonuclease -
HGD79_RS24765 () 89257..89682 + 426 WP_001043843.1 hypothetical protein -
HGD79_RS24770 () 90136..90372 + 237 Protein_88 hypothetical protein -
HGD79_RS24775 () 90427..90726 + 300 WP_001572440.1 hypothetical protein -
HGD79_RS24780 () 91039..93078 - 2040 WP_032192809.1 ATP-dependent helicase -
HGD79_RS24785 () 93075..94061 - 987 WP_001572342.1 hypothetical protein -
HGD79_RS24790 () 94647..95030 + 384 Protein_92 HNH endonuclease -
HGD79_RS24795 () 95092..95295 + 204 WP_001194555.1 hypothetical protein -
HGD79_RS24800 () 95637..96041 - 405 WP_001287392.1 H-NS histone family protein -
HGD79_RS24805 () 96539..96775 - 237 WP_000175476.1 hypothetical protein -
HGD79_RS24810 () 96817..97272 - 456 WP_001572344.1 hypothetical protein -
HGD79_RS24815 () 97332..97997 - 666 WP_000490638.1 hypothetical protein -
HGD79_RS24820 () 98055..98435 - 381 WP_001426317.1 hypothetical protein -
HGD79_RS24825 () 99078..99896 + 819 WP_168848543.1 DUF4942 domain-containing protein -
HGD79_RS24830 () 99893..101098 + 1206 WP_001572381.1 hypothetical protein -
HGD79_RS24835 () 101378..102697 - 1320 WP_000078514.1 DUF1173 family protein -
HGD79_RS24840 () 102720..102887 - 168 WP_000220758.1 hypothetical protein -
HGD79_RS24845 () 102948..104375 - 1428 WP_000833382.1 DNA cytosine methyltransferase -
HGD79_RS24850 () 104590..105105 + 516 WP_001572389.1 thermonuclease family protein -
HGD79_RS24855 () 105108..105947 + 840 WP_168848544.1 hypothetical protein -
HGD79_RS24860 () 106226..106459 + 234 WP_015059496.1 hypothetical protein -
HGD79_RS24865 () 107121..107351 + 231 WP_001572393.1 hypothetical protein -
HGD79_RS24870 () 107688..108149 + 462 WP_000185304.1 hypothetical protein -
HGD79_RS24875 () 108179..108586 + 408 WP_000074418.1 hypothetical protein -
HGD79_RS24880 () 108637..108954 - 318 WP_029698059.1 hypothetical protein -
HGD79_RS24885 () 109331..109681 - 351 WP_031613424.1 hypothetical protein -
HGD79_RS24890 () 109791..111305 - 1515 Protein_112 Tn3-like element TnAs1 family transposase -
HGD79_RS24895 () 111371..112075 + 705 WP_001067855.1 IS6 family transposase IS26 Transposase
HGD79_RS24905 () 112378..113253 + 876 WP_001617865.1 Beta-lactamase CTX-M-97 ARG
HGD79_RS24910 () 113288..113605 + 318 Protein_115 IS5/IS1182 family transposase -
HGD79_RS24915 () 113865..114281 + 417 WP_014839980.1 Glutathione transferase FosA ARG
HGD79_RS24920 () 114286..114804 - 519 WP_014839979.1 hypothetical protein -
HGD79_RS24925 () 114804..115550 - 747 WP_071448054.1 winged helix-turn-helix domain-containing protein -
HGD79_RS24930 () 115556..116260 - 705 WP_001067855.1 IS6 family transposase IS26 Transposase
HGD79_RS24935 () 116374..117150 + 777 WP_001199192.1 SPBc2 prophage-derived aminoglycoside N(3')-acetyltransferase-like protein YokD ARG
HGD79_RS24940 () 117379..118279 + 901 Protein_121 phosphotransferase -
HGD79_RS24945 () 118285..118650 + 366 Protein_122 AAC(3) family N-acetyltransferase -
HGD79_RS24950 () 118701..119453 - 753 WP_000602738.1 IS6 family transposase ISRle7 Transposase
HGD79_RS24955 () 119539..121038 + 1500 Protein_124 Tn3 family transposase TnpA_ISRsp12 Transposase
HGD79_RS24960 () 121114..121824 - 711 WP_057102336.1 arsenical resistance protein ArsH -
HGD79_RS24965 () 121925..122248 - 324 WP_001175593.1 helix-turn-helix domain-containing protein -
HGD79_RS24970 () 122354..123487 + 1134 WP_058650130.1 permease -
HGD79_RS24975 () 123780..124397 - 618 WP_000521603.1 hypothetical protein -
HGD79_RS24980 () 124589..126145 - 1557 WP_000818556.1 AAA family ATPase -
HGD79_RS24985 () 126408..126998 - 591 WP_000194575.1 hypothetical protein -
HGD79_RS24990 () 126998..127255 - 258 WP_000343085.1 hypothetical protein -
HGD79_RS24995 () 127609..129747 + 2139 WP_000350638.1 AAA family ATPase -
HGD79_RS25000 () 129909..130325 - 417 WP_001044768.1 type II toxin-antitoxin system VapC family toxin Toxin Detail
HGD79_RS25005 () 130322..130552 - 231 WP_001261287.1 AbrB/MazE/SpoVT family DNA-binding domain-containing protein Antitoxin Detail
HGD79_RS25010 () 130848..131120 + 273 WP_061602529.1 hypothetical protein -
HGD79_RS25015 () 131167..131871 + 705 WP_001067858.1 IS6 family transposase IS15DI Transposase
HGD79_RS25020 () 131918..132352 - 435 WP_000429836.1 IS607 family transposase ISChh1 Transposase
HGD79_RS25025 () 132424..132774 + 351 WP_001294663.1 mercuric transport protein MerT -
HGD79_RS25030 () 132788..133063 + 276 WP_000732292.1 mercury resistance system periplasmic binding protein MerP -
HGD79_RS25035 () 133099..133521 + 423 WP_001340589.1 organomercurial transporter MerC -
HGD79_RS25040 () 133573..135267 + 1695 WP_000105636.1 mercury(II) reductase -
HGD79_RS25045 () 135285..135647 + 363 WP_001277456.1 mercury resistance co-regulator MerD -
HGD79_RS25050 () 135644..135880 + 237 WP_000993386.1 broad-spectrum mercury transporter MerE -
HGD79_RS25055 () 135916..136584 + 669 WP_001300294.1 putative signaling protein VF
HGD79_RS25060 () 136659..137909 + 1251 Protein_145 transposase -
HGD79_RS25065 () 137974..138678 + 705 WP_001067855.1 IS6 family transposase IS26 Transposase
HGD79_RS25070 () 139128..140357 + 1230 WP_032492336.1 Putative bacilysin exporter BacE ARG
HGD79_RS25075 () 140503..141366 + 864 WP_000612791.1 SDR family oxidoreductase -
HGD79_RS25080 () 141404..141649 + 246 WP_001354008.1 GrpB family protein -
HGD79_RS25085 () 142118..142909 + 792 WP_000034420.1 Dihydropteroate synthase ARG
HGD79_RS25090 () 142912..143187 - 276 WP_109023896.1 hypothetical protein -
HGD79_RS25095 () 143189..143824 + 636 Protein_152 IS256 family transposase IS406 Transposase
HGD79_RS25100 () 144089..144421 - 333 WP_000800531.1 quaternary ammonium compound efflux SMR transporter QacL -
HGD79_RS25105 () 144591..145382 - 792 WP_001206316.1 Streptomycin 3''-adenylyltransferase ARG
HGD79_RS25110 () 145475..146734 - 1260 WP_000095725.1 Bicyclomycin resistance protein ARG
HGD79_RS25115 () 146996..147787 - 792 WP_001206356.1 Streptomycin 3''-adenylyltransferase ARG
HGD79_RS25120 () 147793..148083 - 291 WP_001336345.1 DUF1010 domain-containing protein -
HGD79_RS25125 () 148195..148692 - 498 WP_001083725.1 Dihydrofolate reductase ARG
HGD79_RS25130 () 148837..149808 + 972 WP_119015029.1 IS91 family transposase ISAzo26 Transposase
HGD79_RS25135 () 149842..150546 - 705 WP_001067855.1 Tn3 family transposase TnpA_Tn3 Transposase
HGD79_RS25140 () 150610..151200 + 591 Protein_161 Tn3 family transposase TnpA_Tn3 Transposase
HGD79_RS25145 () 151243..152258 - 1016 Protein_162 IS3 family transposase ISEc36 Transposase
HGD79_RS25150 () 152653..153309 + 657 WP_001516695.1 Pentapeptide repeat protein ARG
HGD79_RS25155 () 153628..154056 - 429 Protein_164 Tn3 family transposase TnpR_ISMex22 Transposase
HGD79_RS25160 () 154089..155480 - 1392 WP_001493761.1 ISKra4 family transposase ISKpn19 Transposase
HGD79_RS25165 () 155517..156089 - 573 WP_001493762.1 Tn3 family transposase TnpR_ISPa40 Transposase
HGD79_RS25170 () 156226..156816 - 591 WP_012477564.1 plasmid pRiA4b ORF-3 family protein -
HGD79_RS25175 () 156923..157627 - 705 WP_001067855.1 IS6 family transposase IS26 Transposase
HGD79_RS25180 () 157688..158506 + 819 Protein_169 transposase -
HGD79_RS25185 () 158564..158806 + 243 WP_000844627.1 transposase -
HGD79_RS25190 () 158838..159488 - 651 WP_063120597.1 tetracycline resistance transcriptional repressor TetR(A) -
HGD79_RS25195 () 159594..160793 + 1200 WP_001493765.1 Tetracycline resistance protein, class C ARG
HGD79_RS25200 () 160825..161709 - 885 WP_000058717.1 EamA family transporter -
HGD79_RS25205 () 161847..162254 - 408 WP_001214976.1 cysteine hydrolase -
HGD79_RS25210 () 162243..163994 + 1752 Protein_175 Tn3 family transposase TnpA_TnAs1 Transposase
HGD79_RS25215 () 164056..165687 - 1632 WP_072656931.1 copper resistance system multicopper oxidase -
HGD79_RS25220 () 165922..166371 + 450 WP_001023231.1 hypothetical protein -
HGD79_RS25225 () 166646..167383 + 738 WP_000287499.1 Murein DD-endopeptidase MepM VF
HGD79_RS25230 () 167417..167614 - 198 WP_000843501.1 DUF2933 domain-containing protein -
HGD79_RS25235 () 167655..170132 - 2478 WP_011251358.1 Ag(+)-translocating P-type ATPase SilP -
HGD79_RS25240 () 170230..170670 - 441 WP_000758221.1 hypothetical protein -
HGD79_RS25245 () 170757..173903 - 3147 WP_011251357.1 Cu(+)/Ag(+) efflux RND transporter permease subunit SilA -
HGD79_RS25250 () 173914..175206 - 1293 WP_001398209.1 Cu(+)/Ag(+) efflux RND transporter periplasmic adaptor subunit SilB -
HGD79_RS25255 () 175320..175673 - 354 WP_001246155.1 cation efflux system protein CusF -
HGD79_RS25260 () 175701..177086 - 1386 WP_063107410.1 Cation efflux system protein CusC VF
HGD79_RS25265 () 177276..177956 + 681 WP_000697965.1 Transcriptional regulatory protein CusR VF
HGD79_RS25270 () 177949..179424 + 1476 WP_000555737.1 copper/silver sensor histidine kinase SilS -
HGD79_RS25275 () 179675..180106 + 432 WP_000790483.1 silver-binding protein SilE -
HGD79_RS25280 () 180250..180600 + 351 WP_000694953.1 DUF305 domain-containing protein -
HGD79_RS25285 () 180988..181896 - 909 WP_001372261.1 HNH endonuclease -
HGD79_RS25290 () 182446..183150 - 705 WP_001067858.1 IS6 family transposase IS15DI Transposase
HGD79_RS25295 () 183525..183782 - 258 WP_042634392.1 hypothetical protein -
HGD79_RS25300 () 183848..184174 - 327 WP_000556022.1 hypothetical protein -
HGD79_RS25305 () 184590..184736 - 147 WP_001426334.1 hypothetical protein -
HGD79_RS25310 () 185017..186267 - 1251 WP_000104323.1 site-specific DNA-methyltransferase -
HGD79_RS25315 () 186436..187044 - 609 WP_000833773.1 hypothetical protein -
HGD79_RS25320 () 187200..187457 - 258 WP_001290637.1 hypothetical protein -
HGD79_RS25325 () 187522..187950 - 429 WP_001133498.1 hypothetical protein -
HGD79_RS25330 () 188045..188422 - 378 WP_000853477.1 hypothetical protein -
HGD79_RS25335 () 188830..190011 - 1182 Protein_200 flavin reductase family protein -
HGD79_RS25340 () 190020..190316 - 297 WP_000581857.1 type II toxin-antitoxin system RelE/ParE family toxin -
HGD79_RS25345 () 190366..190836 - 471 WP_000170638.1 hypothetical protein -
HGD79_RS25350 () 191271..193661 - 2391 WP_000694550.1 Ig-like domain repeat protein -
HGD79_RS25355 () 193734..193901 - 168 WP_000803860.1 hypothetical protein -
HGD79_RS25360 () 194222..194986 + 765 WP_000277497.1 hypothetical protein -
HGD79_RS25365 () 195067..195828 + 762 WP_001167416.1 methyltransferase -
HGD79_RS25370 () 195922..196185 + 264 WP_000149861.1 hypothetical protein -
HGD79_RS25375 () 196230..196817 + 588 WP_000491822.1 hypothetical protein -
HGD79_RS25380 () 197282..199702 + 2421 WP_000681217.1 DUF4165 domain-containing protein -
HGD79_RS25385 () 201700..202788 - 1089 WP_000159529.1 IncHI-type plasmid replication initiator protein RepHI2 -
HGD79_RS25390 () 203896..206352 - 2457 WP_001572417.1 DUF4165 domain-containing protein -
HGD79_RS25395 () 206847..208075 + 1229 WP_152919722.1 IS3 family transposase IS2 Transposase
HGD79_RS25400 () 208091..208687 + 597 WP_046788420.1 hypothetical protein -
HGD79_RS25405 () 208745..209026 - 282 WP_000535422.1 hypothetical protein -
HGD79_RS25410 () 209007..209468 - 462 WP_000502497.1 hypothetical protein -
HGD79_RS25415 () 209517..209693 - 177 WP_000070863.1 hypothetical protein -
HGD79_RS25420 () 209690..211381 - 1692 WP_042634269.1 DUF4942 domain-containing protein -
HGD79_RS25425 () 211629..212792 - 1164 WP_000114859.1 hypothetical protein -
HGD79_RS25430 () 213162..213974 + 813 WP_001121575.1 Dam family site-specific DNA-(adenine-N6)-methyltransferase -
HGD79_RS25435 () 213983..214837 - 855 WP_000970403.1 hypothetical protein -
HGD79_RS25440 () 215072..215788 - 717 WP_046788421.1 zinc finger-like domain-containing protein -
HGD79_RS25445 () 215785..215985 - 201 WP_000594931.1 hypothetical protein -
HGD79_RS25450 () 216003..217058 - 1056 WP_001065779.1 hypothetical protein -

Similar MGE(s)


Similar MGE(s) is defined as other TA-related MGE(s) with Mash distance < 0.01 to this MGE.

Detail Organism MGE type Related TA Genome accession Coordinates Mash
distance