T4SS ID	262
Strain	Cupriavidus metallidurans CH34
Replicon	chromosome [Browse all T4SS(s) in this replicon]
Accession	NC_007973
Location	1378405..1378992; 1448292..1460364
Name	Trb
Function	conjugation
Classification	Type IVA; Type P
This T4SS has been mentioned in a genome sequencing report.

T4SS components

Component	TrbB	TrbC	TrbD	TrbE	TrbF	TrbG	TrbH	TrbI	TrbJ
Number	1	1	1	1	1	1	0	1	1
Component	TrbK	TrbL	TrbN	TraF	TraG
Number	0	1	0	1	1

The information of T4SS components from NC_007973
Region 1: 1378405..1378992
#	Locus tag (Gene)	Coordinates [+/-], size (bp)	Protein GI	Product	Component
1	Rmet_1251 (tnp)	1373579..1374580 [-], 1002	94310195	transposase ISRme5 (copy a, CMGI-2)
2	Rmet_1252	1374596..1375468 [+], 873	94310196	hypothetical protein
3	Rmet_1253	1375516..1375833 [+], 318	94310197	putative dna binding protein
4	Rmet_1254 (repA)	1375860..1376717 [+], 858	94310198	replication initiator and transcription repressor protein
5	Rmet_1255 (parA)	1376971..1377609 [+], 639	94310199	ATPase involved in plasmid partitioning
6	Rmet_1256 (parG)	1377606..1377878 [+], 273	94310200	regulator of plasmid copy number
7	Rmet_1257	1377875..1378408 [+], 534	94310201	hypothetical protein
8	Rmet_1258 (traF)	1378405..1378992 [+], 588	94310202	plasmid conjugal transfer transmembrane protein	TraF
9	Rmet_1259 (virD2)	1379309..1381414 [+], 2106	94310203	relaxase type IV secretory pathway VirD2
10	Rmet_1260	1381452..1381913 [-], 462	94310204	hypothetical protein
11	Rmet_1261	1381974..1383101 [-], 1128	94310205	putative secreted protein
Region 2: 1448292..1460364
#	Locus tag (Gene)	Coordinates [+/-], size (bp)	Protein GI	Product	Component
1	Rmet_1331 (tomA0)	1443460..1443684 [-], 225	94310273	TomA0
2	Rmet_1333 (phaB2)	1444571..1445317 [+], 747	94310275	3-oxoacyl-(acyl-carrier-protein) reductase
3	Rmet_1334 (uspA2)	1445377..1445841 [+], 465	94310276	universal stress protein, UspA family
4	Rmet_1335 (chnA)	1446018..1446806 [+], 789	94310277	cyclohexanol dehydrogenase
5	Rmet_1336 (traR)	1446803..1447765 [+], 963	94310278	LysR family transcriptional regulator
6	Rmet_1337	1447762..1448040 [+], 279	94310279	lipoprotein
7	Rmet_1338	1447986..1448282 [-], 297	94310280	hypothetical protein
8	Rmet_1339 (traG)	1448292..1450286 [+], 1995	94310281	conjugal transfer coupling protein TraG	TraG
9	Rmet_1340	1450283..1450747 [+], 465	94310282	CopG-like DNA-binding helix-turn-helix domain-containing protein
10	Rmet_1341 (trbB)	1450744..1451808 [+], 1065	94310283	P-type conjugative transfer ATPase TrbB	TrbB
11	Rmet_1342 (trbC)	1451805..1452188 [+], 384	94310284	conjugal transfer TrbC transmembrane protein	TrbC
12	Rmet_1343 (trbD)	1452185..1452457 [+], 273	94310285	conjugal transfer TrbD transmembrane protein	TrbD
13	Rmet_1344 (trbE)	1452470..1454929 [+], 2460	94310286	conjugal transfer ATPase TrbE	TrbE
14	Rmet_1345 (trbJ)	1454926..1455663 [+], 738	94310287	conjugal transfer protein TrbJ	TrbJ
15	Rmet_1346	1455676..1455996 [+], 321	94310288	lipoprotein
16	Rmet_1347 (trbL)	1455993..1457372 [+], 1380	94310289	conjugal transfer protein TrbL	TrbL
17	Rmet_1348 (trbF)	1457397..1458101 [+], 705	94310290	conjugal transfer protein TrbF	TrbF
18	Rmet_1349 (trbG)	1458098..1459090 [+], 993	94310291	conjugal transfer protein TrbG/VirB9/CagX putative exported protein	TrbG
19	Rmet_1350 (trbI)	1459093..1460364 [+], 1272	94310292	type IV secretion system transmembrane protein	TrbI
20	Rmet_1351	1460361..1460606 [+], 246	94310293	hypothetical protein
21	Rmet_1352 (comL)	1460941..1461777 [-], 837	94310294	DNA uptake lipoprotein
22	Rmet_1353 (rluD)	1461789..1463000 [+], 1212	94310295	23S rRNA pseudouridine synthase
23	Rmet_1354	1462987..1463778 [+], 792	94310296	hypothetical protein
24	Rmet_1355	1463804..1464031 [-], 228	94310297	putative transmembrane protein

Genes in the 5-Kb flanking regions if available, or non-essential genes in the T4SS gene cluster if any.

Download FASTA format files

Proteins        Genes

This T4SS is encoded by the integrative and conjugative element (ICE) CMGI-2 and is thought to contribute to the transfer of this element. (click on the ICE name to view the detail ICE informaion in ICEberg, an ICE database)

(1) Janssen PJ; Van Houdt R; Moors H; Monsieurs P; Morin N; Michaux A; Benotmane MA; Leys N; Vallaeys T; Lapidus A; Monchy S; Medigue C; Taghavi S; McCorkle S; Dunn J; van der Lelie D; Mergeay M (2010). The complete genome sequence of Cupriavidus metallidurans strain CH34, a master survivalist in harsh and anthropogenic environments. PLoS One. 5(5):e10433. [PudMed:20463976]